Universities in the USA Offering Full Scholarships- Application links, Requirements, How To Apply
| Institution | Program | Program Link |
| Albert Einstein College of Medicine | Biomedical Sciences | https://www.einstein.yu.edu/education/phd/prospective-students/admissions.aspx |
| Auburn University | Biological Sciences | http://www.auburn.edu/cosam/departments/biology/grad/index.htm |
| Baylor College of Medicine | Graduate School of Biomedical Sciences | https://www.bcm.edu/education/schools/graduate-school-of-biomedical-sciences |
| Boston University | Molecular Biology, Cell Biology, & Biochemistry | http://www.bu.edu/mcbb/admissions/ |
| Boston University | Program in Biomedical Science | http://www.bumc.bu.edu/gms/pibs/ |
| Brandeis University | Molecular and Cell Biology | http://www.bio.brandeis.edu/grad/mcb/mcb_phd_rev.html |
| Brandeis University | Neuroscience | http://www.bio.brandeis.edu/grad/neuro/neuro_phd.html |
| Brown University | Biomedical Engineering | https://www.brown.edu/graduateprograms/biomedical-engineering-phd |
| Brown University | Biotechnology | https://www.brown.edu/graduateprograms/biomed-biotechnology-phd |
| Brown University | Computational Biology | https://www.brown.edu/graduateprograms/computational-biology-phd |
| Brown University | Ecology and Evolutionary Biology | https://www.brown.edu/graduateprograms/biomed-ecology-and-evolutionary-biology-phd |
| Brown University | Molecular Biology, Cell Biology and Biochemistry | https://www.brown.edu/graduateprograms/biomed-molecular-biology-cell-biology-and-biochemistry-phd |
| Brown University | Molecular Pharmacology and Physiology | https://www.brown.edu/graduateprograms/biomed-molecular-pharmacology-and-physiology-phd |
| Brown University | Neuroscience | https://www.brown.edu/graduateprograms/biomed-neuroscience-phd |
| Brown University | Pathobiology | https://www.brown.edu/graduateprograms/pathobiology-phd |
| Binghamton University – SUNY | Behavioral Neuroscience | https://www.binghamton.edu/psychology/graduate/behavioral-neuroscience/index.html |
| Binghamton University – SUNY | Clinical Psychology | https://www.binghamton.edu/psychology/graduate/clinical-psychology/index.html |
| Binghamton University – SUNY | Biological Sciences | https://www.binghamton.edu/biology/graduate/index.html |
| California State University Dominguez Hills | Biology | https://www.csudh.edu/gsr/graduate-studies/graduate-admission/# |
| California State University Sacramento | Master’s of Stem Cel Biology | https://www.csus.edu/college/natural-sciences-mathematics/biology/stem-cell-program.html |
| Caltech | Biochemistry and Molecular Biophysics | https://cce.caltech.edu/graduate/graduate-biochemistry-and-molecular-biophysics |
| Carnegie Mellon University | Biological Sciences | https://www.cmu.edu/bio/graduate/phd_program/admissions.html |
| Case Western Reserve University | Biomedical Sciences Training Program | https://case.edu/med/BSTP/apply.html |
| City University of New York | Biology | https://www.gc.cuny.edu/Prospective-Current-Students/Prospective-Students/Admissions#! |
| Clemson University | Biological Sciences | https://www.clemson.edu/science/departments/biosci/academics/graduate/plan-study-biosci.html |
| Clemson University | Environmental Toxicology | https://www.clemson.edu/science/departments/biosci/academics/graduate/plan-study-entox.html |
| Clemson University | Microbiology | https://www.clemson.edu/science/departments/biosci/academics/graduate/plan-study-micro.html |
| Cold Spring Harbor Laboratory | PhD Program | https://www.cshl.edu/education/phd-program/how-to-apply/ |
| Colorado State University | Biochemistry and Molecular Biology | https://www.bmb.colostate.edu/ |
| Columbia University | Ecology, Evolution, and Environmental Biology | http://e3b.columbia.edu/ |
| Columbia University Medical School | Biomedical Sciences Programs | https://www.gsas.cuimc.columbia.edu/applying |
| Cornell University | Neurobiology & Behavior | https://nbb.cornell.edu/ |
| Cornell University | Biochemistry, Molecular, and Cell Biology | https://bmcb.cornell.edu/ |
| Cornell University | Biomedical Engineering | https://www.bme.cornell.edu/bme/programs/graduate-programs |
| Cornell University | Entomology | https://entomology.cals.cornell.edu/graduate/ |
| Cornell University | Genetics, Genomics, and Development | https://gendev.cornell.edu/ |
| Cornell University | Plant Biology | https://plantbio.cals.cornell.edu/academics/graduate/ |
| Cornell University | Plant Pathology | https://pppmb.cals.cornell.edu/graduate/ |
| Cornell University | Biological and Biomedical sciences | https://www.vet.cornell.edu/education/biomedical-biological-sciences-bbs-phd-program |
| Cornell University | Ecology & Evolutionary Biology | https://ecologyandevolution.cornell.edu/ |
| Dartmouth | Molecular & Cell Biology | https://graduate.dartmouth.edu/mcb/ |
| Dartmouth | Program in Experimental & Molecular Medicine | https://graduate.dartmouth.edu/pemm/ |
| DePaul University | Biological Sciences (MS) | https://csh.depaul.edu/academics/biological-sciences/graduate/biological-sciences-ms/Pages/default.aspx |
| Drexel University | Neuroscience (MS or PhD) | https://drexel.edu/medicine/academics/graduate-school/neuroscience/ |
| Drexel University | Pharmacology and Physiology | https://drexel.edu/medicine/academics/graduate-school/pharmacology-physiology/ |
| Duke University | Neurobiology | https://www.neuro.duke.edu/ |
| Duke University | Biochemistry | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Cell & Molecular Biology | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Developmental & Stem Cell Biology | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Immunology | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Molecular Cancer Biology | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Molecular Genetics and Microbiology | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Pharmacology | https://gradschool.duke.edu/admissions/apply/gre-scores |
| Duke University | Biology | https://biology.duke.edu/graduate |
| Duke University | Genetics and Genomics | https://upg.duke.edu/ |
| Eastern Virginia Medical School | Biomedical Sciences | https://www.evms.edu/education/doctoral_programs/biomedical_sciences/ |
| Emory University | Chemistry | http://chemistry.emory.edu/home/graduate/apply.html |
| Emory University | GDBBS – Biochemistry, Cell and Developmental Biology | http://www.biomed.emory.edu/bcdb |
| Emory University | GDBBS – Cancer Biology | http://www.biomed.emory.edu/cb |
| Emory University | GDBBS – Genetics and Molecular Biology | http://www.biomed.emory.edu/gmb |
| Emory University | GDBBS – Immunology and Molecular Pathogenesis | http://www.biomed.emory.edu/imp |
| Emory University | GDBBS – Microbiology and Molecular Genetics | http://www.biomed.emory.edu/mmg |
| Emory University | GDBBS – Molecular and Systems Pharmacology | http://www.biomed.emory.edu/msp |
| Emory University | GDBBS – Neuroscience | http://www.biomed.emory.edu/ns |
| Emory University | GDBBS – Population Biology, Ecology, and Evolution | http://www.biomed.emory.edu/pbee |
| Florida State University | Biological Sciences | https://www.bio.fsu.edu/ |
| Florida State University | Biomedical Sciences | https://med.fsu.edu/phd/home |
| Florida State University | Neuroscience | https://neuro.fsu.edu/gr/admission |
| Georgetown University Biomedical Graduate Education | Pharmacology and Physiology | https://pharmacology.georgetown.edu/program/ |
| Georgetown University Biomedical Graduate Education | Interdisciplinary Program in Neuroscience | https://neuroscience.georgetown.edu/ |
| Georgetown University Biomedical Graduate Education | Tumor Biology | https://tumorbiology.georgetown.edu/ |
| Georgetown University Biomedical Graduate Education | Biomedical masters degree programs | https://biomedicalprograms.georgetown.edu/masters-programs/ |
| Georgetown University | Biology | https://biology.georgetown.edu/graduate/ |
| George Washington University | IBS – Cancer Biology program | https://smhs.gwu.edu/ibs/phd-programs-ibs/cancer-biology-phd-program |
| George Washington University | IBS – Genomics & Bioinformatics | https://smhs.gwu.edu/ibs/phd-programs-ibs/genomics-bioinformatics-phd-program |
| George Washington University | IBS – Microbiology & Immunology | https://smhs.gwu.edu/ibs/program/microbio-immunology |
| George Washington University | IBS – Neuroscience | https://smhs.gwu.edu/ibs/program/molecular-medicine/neurosciences |
| George Washington University | IBS – Pharmacology & Physiology | https://smhs.gwu.edu/ibs/program/molecular-medicine/pharma-physio |
| Gerstner Sloan Kettering | Biomedical Sciences | https://www.sloankettering.edu/gerstner/admissions/apply |
| Harvard University | Biological and Biomedical Sciences | http://www.hms.harvard.edu/dms/bbs/ |
| Harvard University | Biological Sciences in Dental Medicine | https://gsas.harvard.edu/programs-of-study/all/biological-sciences-dental-medicine |
| Harvard University | Biological Sciences in Public Health | https://www.hsph.harvard.edu/biological-sciences/ |
| Harvard University | Biological Sciences in Public Health | https://www.hsph.harvard.edu/biological-sciences/ |
| Harvard University | Biophysics | https://gsas.harvard.edu/programs-of-study/all/biophysics |
| Harvard University | Chemical Biology | http://chembiophd.harvard.edu/ |
| Harvard University | Immunology | http://dms.hms.harvard.edu/immunology/index.html |
| Harvard University | Molecular and Cellular Biology | https://gsas.harvard.edu/programs-of-study/all/molecular-and-cellular-biology |
| Harvard University | Neuroscience | http://dms.hms.harvard.edu/neuroscience/index.html |
| Harvard University | Organismic & Evolutionary Biology | https://gsas.harvard.edu/programs-of-study/all/organismic-and-evolutionary-biology |
| Harvard University | Systems Biology | http://sysbiophd.harvard.edu/prospective-students/admissions |
| Harvard University | Virology | http://dms.hms.harvard.edu/virology/index.html |
| Icahn School of Medicine at Mount Sinai | Biomedical Science, Neuroscience | http://icahn.mssm.edu/education/phd |
| Illinois State University | Biological Sciences (PhD) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Neuroscience and Physiology) (PhD) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Molecular and Cellular Biology) (PhD) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Behavior,Ecology,Evolution and Systematics) (PhD) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (MS) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Behavior,Ecology,Evolution and Systematics) (MS) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Conservation Biology) (MS) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Biomathematics) (MS) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Bioenergy) (MS) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Biotechnology) (MS) | https://biology.illinoisstate.edu/graduate/ |
| Illinois State University | Biological Sciences (Neuroscience and Physiology) (MS) | https://biology.illinoisstate.edu/graduate/ |
| Indiana University | Biology | https://www.bio.indiana.edu/ |
| Indiana University | Cell, Molecular and Cancer Biology | https://medicine.iu.edu/campuses/bloomington/graduate-programs/cell-molecular-and-cancer-biology-phd/ |
| Indiana University | Neuroscience | https://neuroscience.indiana.edu/academics/how-to-apply/index.html |
| Indiana University School of Medicine | Biomedical | https://medicine.iu.edu/education/graduate-degrees/phd/admissions/ |
| Iowa State University | Ecology and Evolutionary Biology | https://www.eeb.iastate.edu/admissions |
| Johns Hopkins School of Medicine | Biochemistry, Cellular & Molecular Biology | http://bcmb.bs.jhmi.edu/ |
| Johns Hopkins School of Medicine | Biological Chemistry | http://biolchem.bs.jhmi.edu/Pages/application-information.aspx |
| Johns Hopkins School of Medicine | Cellular and Molecular Medicine Program | http://cmm.jhmi.edu/index.php/application-information/ |
| Johns Hopkins School of Medicine | Immunology | https://gradimmunology.med.som.jhmi.edu/ |
| Johns Hopkins School of Medicine | Molecular Biophysics | https://pmb.jhu.edu/admissions/ |
| Johns Hopkins School of Medicine | Neuroscience | http://neuroscience.jhu.edu/graduate/apply |
| Johns Hopkins School of Medicine | Pathobiology | http://pathology.jhu.edu/department/training/pathbio_ProgramOverview.cfm |
| Johns Hopkins School of Medicine | Pharmacology and Molecular Sciences | http://www.hopkinsmedicine.org/pharmacology_molecular_sciences |
| Johns Hopkins Bloomberg School of Public Health | Biochemistry and Molecular Biology | http://www.jhu-bmb-phd.org/ |
| Johns Hopkins University | Biology (MS) | https://bio.jhu.edu/undergraduate/masters-program/ |
| Johns Hopkins University | Cell, Molecular, Developmental Biology, & Biophysics | https://cmdb.jhu.edu/ |
| Johns Hopkins University | Cellular and Molecular Physiology | http://physiology.bs.jhmi.edu/academics/ |
| Johns Hopkins University | Human Genetics and Molecular Biology | https://igm.jhmi.edu/humangenetics/welcome |
| Johns Hopkins University | Molecular Biophysics | http://pmcb.jhu.edu/ |
| Kansas State University | Biology | https://www.k-state.edu/biology/ |
| Massachusetts Institute of Technology | Biological Engineering | http://be.mit.edu/academic-programs/prospective-grad/graduate-application |
| Massachusetts Institute of Technology | Biology | https://biology.mit.edu/graduate/prospective-students/application-process/ |
| Massachusetts Institute of Technology | Brain and Cognitive Science | https://bcs.mit.edu/academic-program/graduate/graduate-admissions |
| Mayo Clinic College of Medicine | Biomedical Sciences | https://college.mayo.edu/academics/biomedical-research-training/admissions-and-benefits/ |
| Medical College of Wisconsin | Interdisciplinary Program in Biomedical Sciences | www.mcw.edu/idp |
| Medical College of Wisconsin | Neuroscience | www.mcw.edu/ndp |
| Medical College of Wisconsin | Pharmacology & Toxicology | www.mcw.edu/pharmtoxphd |
| Medical University of South Carolina | Biomedical Sciences | www.musc.edu/grad |
| Michigan Tech | Biological Sciences | http://www.mtu.edu/gradschool/programs/degrees/biological-sciences/ |
| New York Medical College | Basic Medical Sciences | https://www.nymc.edu/graduate-school-of-basic-medical-sciences-gsbms/admissions–financial-aid/admission-requirements/ |
| New York University | Biology | https://as.nyu.edu/biology/programs/phd.html |
| New York University Sackler Institute | Biomedical Sciences | https://med.nyu.edu/research/sackler-institute-graduate-biomedical-sciences/ |
| NIH Oxford-Cambridge | Scholars Program | https://oxcam.gpp.nih.gov/ |
| Northwestern University | Neuroscience | https://www.nuin.northwestern.edu/application-process-2/ |
| Northwestern University | Biological Sciences | https://www.ibis.northwestern.edu/graduate/prospective/ |
| Northwestern University | Life Sciences | http://www.feinberg.northwestern.edu/sites/dgp/prospective-students/phd-program/admissions.html |
| Ohio State University | Biochemistry | https://osbp.osu.edu/admissions |
| Ohio State University | Evolution, Ecology and Organismal Biology | https://eeob.osu.edu/grad/graduate-program |
| Ohio State University | Microbiology | https://microbiology.osu.edu/gradstudents/admissions |
| Ohio State University | Molecular Genetics | https://molgen.osu.edu/program-admissions-and-aid |
| Ohio State University | Chemical and Biomolecular Engineering | https://cbe.osu.edu/phd |
| Oregon State University | Biochemistry and Biophysics | https://biochem.oregonstate.edu/ |
| Penn State College of Medicine | Anatomy | https://med.psu.edu/anatomy-phd |
| Penn State College of Medicine | Biomedical Sciences | https://med.psu.edu/biomedical-sciences-phd |
| Penn State College of Medicine | Neuroscience | https://med.psu.edu/neuroscience-phd |
| Penn State University | Biochemistry, Microbiology, and Molecular Biology | https://science.psu.edu/bmb |
| Penn State University | Bioinformatics and Genomics | https://www.huck.psu.edu/graduate-programs/bioinformatics-and-genomics/ |
| Penn State University | Biology | https://science.psu.edu/bio |
| Princeton University | Ecology and Evolutionary Biology | https://eeb.princeton.edu/ |
| Princeton University | Molecular Biology | https://molbio.princeton.edu/ |
| Princeton University | Neuroscience | http://pni.princeton.edu/ |
| Purdue University | Biological Sciences | https://www.bio.purdue.edu/ |
| Purdue University | Chemistry | https://www.chem.purdue.edu/faq/index.php#gre |
| Richard Gilder Graduate School- American Museum of Natural History | Comparative Biology | https://www.amnh.org/research/richard-gilder-graduate-school/apply |
| Rockefeller University | Graduate Program | https://graduateapplication.rockefeller.edu/ |
| Roswell Park Cancer Institute | Cancer Sciences | https://www.roswellpark.org/education/phd-program |
| Rutgers University | Biochemistry (Molecular Biosciences) | https://molbiosci.rutgers.edu/ |
| Rutgers University | Cell and Developmental Biology (Molecular Biosciences) | https://molbiosci.rutgers.edu/ |
| Rutgers University | Microbiology & Molecular Genetics (Molecular Biosciences) | https://molbiosci.rutgers.edu/ |
| Rutgers University | Neuroscience | http://rwjms.rutgers.edu/gsbs/programs/neuroscience/index.html |
| Rutgers University | Pharmacology, Cellular & Molecular (Molecular Biosciences) | https://molbiosci.rutgers.edu/ |
| Rutgers University | Physiology & Integrative Biology (Molecular Biosciences) | https://molbiosci.rutgers.edu/ |
| Rutgers University | Toxicology | https://eohsi.rutgers.edu/divisions/toxicology/ |
| Rutgers University (Newark) | Cell Biology, Neuroscience and Physiology (Newark) | http://njms.rutgers.edu/sgs/prospective_students/multidisciplinary_cbnp2.php |
| Rutgers University (Newark) | Immunity and Inflammation (Newark) | http://njms.rutgers.edu/sgs/current_students/multidisciplinary_i3.php |
| Rutgers University (Newark) | Molecular Biology Genetics and Cancer (Newark) | http://njms.rutgers.edu/sgs/prospective_students/multidisciplinary_mbgc.php |
| San Francisco State University | Biology (Master’s) | http://biology.sfsu.edu/content/applying-sfsu-biology-graduate-programs |
| Scripps Research Institute | Skaggs Graduate School | https://education.scripps.edu/graduate/admissions/faqs/ |
| Stanford University | Bioengineering | https://bioengineering.stanford.edu/ |
| Stanford University | Biosciences | https://biosciences.stanford.edu/prospective-students/how-to-apply/ |
| Stanford University | Chemistry | https://chemistry.stanford.edu/academics/phd-program/admissions/how-apply |
| Stowers Institute for Medical Research | Graduate Program | http://www.stowers.org/gradschool/admissions |
| Texas A&M University | Biology | https://www.bio.tamu.edu/ |
| Texas A&M University | Ecology and Evolutionary Biology | https://eeb.tamu.edu/ |
| Texas A&M University | Genetics | https://genetics.tamu.edu/ |
| Thomas Jefferson University | Graduate School of Biomedical Sciences | https://www.jefferson.edu/university/life-sciences.html |
| Tufts University | Conservation Medicine (MS) | https://grad.vet.tufts.edu/ms-conservation-medicine/ |
| University of Alabama Birmingham | Biomedical Sciences | http://www.uab.edu/gbs/home/admissions |
| University of Arizona | Biological and Biomedical Sciences | https://abbs.arizona.edu/ |
| University of Arizona | School of Plant Sciences | https://cals.arizona.edu/spls/home |
| University of Buffalo | Program in Biomedical Science (PPBS) | http://medicine.buffalo.edu/phdprogram.html |
| University of Buffalo | Microbiology and Immunology | http://medicine.buffalo.edu/departments/micro/education/phd.html |
| University of California Berkeley | Biophysics | https://biophysics.berkeley.edu/admissions/ |
| University of California Berkeley | Environmental Science, Policy, and Management | https://ourenvironment.berkeley.edu/graduate-programs/faq |
| University of California Berkeley | Integrative Biology (IB) | http://ib.berkeley.edu/ |
| University of California Berkeley | Microbiology | http://plantandmicrobiology.berkeley.edu/graduate-program-microbiology |
| University of California Berkeley | Molecular and Cell Biology | https://mcb.berkeley.edu/grad/prospective-graduates/admissions-and-recruitment |
| University of California Berkeley | Neuroscience | https://neuroscience.berkeley.edu/ph-d-program/ |
| University of California Berkeley | Plant Biology | http://plantandmicrobiology.berkeley.edu/plant-biology-0 |
| University of California Berkeley | Bioengineering (joint with UC San Francisco) | https://bioegrad.berkeley.edu/prospectivegrads/admissions |
| University of California Davis | Biochemistry, Molecular, Cellular and Developmental Biology | http://bmcdb.ucdavis.edu/admissions/index.html |
| University of California Davis | Ecology | https://ecology.ucdavis.edu/ |
| University of California Davis | Medical Microbiology & Immunology | https://health.ucdavis.edu/medmicro/ |
| University of California Davis | Molecular, Cellular, and Integrative Physiology | https://mcip.ucdavis.edu/ |
| University of California Davis | Nutritional Biology | https://ggnb.ucdavis.edu/ |
| University of California Davis | Plant Biology | https://pbi.ucdavis.edu/about |
| University of California Davis | Plant Pathology | http://plantpathology.ucdavis.edu/gradindex.cfm |
| University of California Davis | Population Biology | https://cpb.ucdavis.edu/ |
| University of California Davis | Horticulture and Agronomy | https://ggha.ucdavis.edu/ |
| University of California Davis | Integrative Pathobiology | https://integrativepath.vetmed.ucdavis.edu/ |
| University of California Davis | Integrative Genetics and Genomics | https://igg.ucdavis.edu/ |
| University of California Davis | Population Biology | https://pbg.ucdavis.edu/ |
| University of California Davis | Neuroscience | https://grad.neuroscience.ucdavis.edu/apply |
| University of California Irvine | Ecology and Evolutionary Biology | https://ecoevo.bio.uci.edu/graduates/admissions/ |
| University of California Los Angeles | Biochemistry, Molecular, and Structural Biology | https://bmsb.chem.ucla.edu/how-apply |
| University of California Los Angeles | Bioscience (Varies by program) | http://bioscience.ucla.edu/application-procedure |
| University of California Los Angeles | Ecology and Evoltionary Biology | https://www.eeb.ucla.edu/graduate/ |
| University of California Merced | Quantitative and Systems Biology | http://qsb.ucmerced.edu/ |
| University of California San Diego | Biomedical Sciences | http://biomedsci.ucsd.edu/how-to-apply/index.html |
| University of California San Francisco | Biomedical Sciences | https://bms.ucsf.edu/ |
| University of California San Francisco | Chemistry and Chemical Biology | https://ccb.ucsf.edu/admissions |
| University of California San Francisco | Developmental and Stem Cell Biology | https://dscb.ucsf.edu/ |
| University of California San Francisco | Integrative Program in Quantitative Biology | https://ipqb.ucsf.edu/ |
| University of California San Francisco | Neuroscience | http://neurograd.ucsf.edu/about-neuroscience-graduate-program |
| University of California San Francisco | Oral and Craniofacial Sciences | https://dentistry.ucsf.edu/programs/oral-cranio-phd |
| University of California San Francisco | Pharmaceutical Sciences and Pharmacogenomics | https://pspg.ucsf.edu/ |
| University of California San Francisco | Rehabilitation Science | https://ptrehab.ucsf.edu/rehabilitation-science-phd |
| University of California San Francisco | Tetrad | https://tetrad.ucsf.edu/admissions |
| University of California Santa Barbara | Biomolecular Science and Engineering | https://www.bmse.ucsb.edu/ |
| University of California Santa Barbara | Molecular, Cellular, and Developmental Biology | https://www.mcdb.ucsb.edu/academics/graduate |
| University of California Santa Cruz | Biomolecular Engineering and Bioinformatics | https://pbse.ucsc.edu/index.html |
| University of California Santa Cruz | Chemical Biology, Biochemistry and Biophysics | https://pbse.ucsc.edu/index.html |
| University of California Santa Cruz | Microbiology and Environmental Toxicology | https://pbse.ucsc.edu/index.html |
| University of California Santa Cruz | Molecular, Cell, and Developmental Biology | https://pbse.ucsc.edu/index.html |
| University of Chicago | Biosciences Programs | https://biosciences.uchicago.edu/programs |
| University of Chicago | Ecology and Evolution | https://pondside.uchicago.edu/ee/index.html |
| University of Colorado Anschutz Medical Campus | Biomedical Sciences | https://www.ucdenver.edu/academics/colleges/Graduate-School/academic-programs/Biomedical/Pages/home.aspx |
| University of Colorado Anschutz Medical Campus | Cancer Biology | https://www.ucdenver.edu/academics/colleges/Graduate-School/academic-programs/cancerbiology/Pages/home.aspx |
| University of Colorado Anschutz Medical Campus | Cell Biology, Stem Cells and Development | https://www.ucdenver.edu/academics/colleges/medicalschool/programs/CSD/Pages/home.aspx |
| University of Colorado Anschutz Medical Campus | Immunology | https://medschool.cuanschutz.edu/immunology-and-microbiology |
| University of Colorado Anschutz Medical Campus | Microbiology | https://medschool.cuanschutz.edu/departments-centers-institutes |
| University of Colorado Anschutz Medical Campus | Molecular Biology | https://www.ucdenver.edu/academics/colleges/medicalschool/programs/Molbio/Pages/home.aspx |
| University of Colorado Anschutz Medical Campus | Pharmaceutical Sciences | https://www.ucdenver.edu/academics/colleges/pharmacy/AcademicPrograms/PhDPrograms/PhDPharmaceuticalSciences/Pages/PhDPharmaceuticalSciences.aspx |
| University of Colorado Anschutz Medical Campus | Pharmacology | https://medschool.cuanschutz.edu/pharmacology |
| University of Colorado Anschutz Medical Campus | Toxicology | https://www.ucdenver.edu/ACADEMICS/COLLEGES/PHARMACY/ACADEMICPROGRAMS/PHDPROGRAMS/PHDTOXICOLOGY/Pages/PhDToxicology.aspx |
| University of Colorado at Boulder | Molecular, Cellular & Developmental Biology | https://www.colorado.edu/mcdb/graduate/application-details-info |
| University of Colorado at Boulder | Ecology and Evolutionary Biology | https://www.colorado.edu/ebio/ |
| University of Connecticut | Molecular and Cell Biology | https://mcb.uconn.edu/application-information-mcb-ph-d-and-ms-graduate-programs-2/ |
| University of Denver | Cell and Molecular Biology (MS & PhD) | https://www.du.edu/academics/admission-steps/biological-sciences-cell-and-molecular-biology-phd-2021-2022 |
| University of Denver | Ecology and Evolutionary Biology (MS & PhD) | https://www.du.edu/academics/admission-steps/biological-sciences-biology-ecology-and-evolution-phd-2021-2022 |
| University of Georgia | Cellular Biology (MS) | https://cellbio.uga.edu/graduate-program |
| University of Georgia | Integrated Life Sciences | https://ils.uga.edu/ |
| University of Illinois at Chicago | Pharmaceutical Sciences Program | https://pharmacy.uic.edu/programs/graduate-programs/phd-programs/phd-in-pharmaceutical-sciences/ |
| University of Illinois at Urbana-Champaign | Chemistry | http://chemistry.illinois.edu/admissions/graduate-admissions |
| University of Illinois at Urbana-Champaign | Molecular and Cellular Biology | https://mcb.illinois.edu/graduate/gradprospect/ |
| University of Iowa | Biomedical Science | https://medicine.uiowa.edu/biomed |
| University of Iowa | Biomedical Engineering Program | https://bme.engineering.uiowa.edu/graduate/bme-graduate-program-phd |
| University of Iowa | Neuroscience | https://neuroscience.grad.uiowa.edu/ |
| University of Kansas | Ecology and Evolutionary Biology | https://eeb.ku.edu/how-apply |
| University of Kansas | Molecular Biosciences | https://molecularbiosciences.ku.edu/ |
| University of Kansas | Bioengineering | http://bio.engr.ku.edu/ |
| University of Kansas Medical Center | Interdisciplinary Graduate Program in Biomedical Sciences | http://www.kumc.edu/igpbs.html |
| University of Kentucky | Biology | https://bio.as.uky.edu/grad-program |
| University of Louisville | Biomedical Sciences-IPIBS | http://louisville.edu/medicine/ipibs/for-students/application-information/application-process |
| University of Maine | School of Biology and Ecology | https://sbe.umaine.edu/ |
| University of Maryland Baltimore | Varies by program (see link) | https://lifesciences.umaryland.edu/prospective/Admissions/ |
| University of Maryland Baltimore | Chemical Engineering | https://cbee.umbc.edu/academics/graduate-application/ |
| University of Massachusetts Amherst | Environmental Conservation | https://eco.umass.edu/ |
| University of Massachusetts Amherst | Molecular and Cellular Biology | http://gpls.cns.umass.edu/mcb |
| University of Massachusetts Amherst | Neuroscience and Behavior | http://gpls.cns.umass.edu/nsb |
| University of Massachusetts Amherst | Organismic & Evolutionary Biology | https://gpls.cns.umass.edu/oeb |
| University of Massachusetts Amherst | Plant Biology | http://gpls.cns.umass.edu/pb |
| University of Massachusetts Medical School | Biomedical Sciences | https://www.umassmed.edu/gsbs/ |
| University of Memphis | Biological Sciences | https://www.memphis.edu/biology/graduate/ |
| University of Miami | Biology | https://biology.as.miami.edu/graduate/index.html |
| University of Michigan | Biophysics | https://lsa.umich.edu/biophysics/graduates/admissions.html |
| University of Michigan | Chemical Biology | http://www.chembio.umich.edu/ |
| University of Michigan | Ecology and Evolutionary Biology | https://lsa.umich.edu/eeb |
| University of Michigan | Program in Biomedical Science | https://medicine.umich.edu/medschool/education/phd-programs/phd-admissions |
| University of Minnesota | Ecology and Evolutionary Biology | https://cbs.umn.edu/academics/departments/eeb/graduate/graduate-admissions |
| University of Minnesota | Biochemistry, Molecular Biology & Biophysics | https://www.med.umn.edu/education-training/graduate-programs |
| University of Minnesota | Integrative Biology and Physiology | https://www.physiology.umn.edu/degrees-and-programs/graduate-program |
| University of Minnesota | Microbiology, Immunology and Cancer Biology | http://www.micab.umn.edu/ |
| University of Minnesota | Molecular, Cellular, Developmental Biology & Genetics | https://www.med.umn.edu/education-training/graduate-programs |
| University of Minnesota | Neuroscience | https://med.umn.edu/neuroscience |
| University of Minnesota | Pharmacology | https://www.med.umn.edu/education-training/graduate-programs |
| University of Minnesota | Plant and Microbial Biology | https://cbs.umn.edu/academics/departments/pmb/graduate-education/admissions |
| University of Missouri | Division of Biological Sciences | http://biology.missouri.edu/graduate-studies/apply/ |
| University of Montana | Organismal Biology, Ecology, and Evolution | http://hs.umt.edu/dbs/grad-programs/OBE/default.php |
| University of Nebraska Medical Center | Interdisciplinary Graduate Program in Biomedical Sciences | https://www.unmc.edu/gradstudies/programs/phd/interdisciplinary-biomedical-sciences.html |
| University of Nevada, Reno’s | Integrative Neuroscience Program | https://www.unr.edu/neuroscience/graduate-program |
| University of New Mexico | Biology | http://biology.unm.edu/graduate/graduate-admissions.shtml |
| University of North Carolina at Chapel Hill | Biological and Biomedical Sciences Program | https://bbsp.unc.edu/ |
| University of North Carolina at Chapel Hill | Biology (incl QBio and EEB) | https://bio.unc.edu/graduate/ |
| University of Notre damme | Biological Sciences | https://biology.nd.edu/faculty-research/areas-of-research/ |
| University of Oklahoma | Microbiology (MS and PhD) | http://www.ou.edu/admissions/academics/arts-sciences/microbiology |
| University of Oklahoma | Microbiology and Plant Biology (MS and PhD) | ou.edu/cas/mpbio |
| University of Oklahoma Health Sciences | Graduate Program in Biomedical Sciences | http://graduate.ouhsc.edu/GraduatePrograms/PhDPrograms/GraduatePrograminBiomedicalSciences.aspx |
| University of Oregon | Biology | https://biology.uoregon.edu/graduate-studies/apply/ |
| University of Oregon | Chemistry and Biochemistry | https://chemistry.uoregon.edu/ |
| University of Pennsylvania | Biology | https://www.bio.upenn.edu/graduate/ |
| University of Pennsylvania | Biomedical Graduate Studies | http://www.med.upenn.edu/bgs/ |
| University of Pittsburgh | Biological Sciences | http://www.biology.pitt.edu/graduate/how-apply |
| University of Pittsburgh | Biomedical Informatics | https://www.dbmi.pitt.edu/content/doctoral-degree-0 |
| University of Pittsburgh | Cell Biology and Molecular Physiology | http://www.gradbiomed.pitt.edu/programs/cell-biology-and-molecular-physiology |
| University of Pittsburgh | Cellular and Molecular Pathology | http://www.gradbiomed.pitt.edu/programs/cellular-molecular-pathology |
| University of Pittsburgh | Clinical and Translational Science | https://www.icre.pitt.edu/ClinTransPhD/index.html |
| University of Pittsburgh | Computational Biology | http://www.compbio.cmu.edu/ |
| University of Pittsburgh | Integrative Systems Biology | https://www.isb.pitt.edu/ |
| University of Pittsburgh | Microbiology and Immunology | http://pmi.pitt.edu/ |
| University of Pittsburgh | Molecular Biophysics & Structural Biology | http://www.mbsb.pitt.edu/ |
| University of Pittsburgh | Molecular Genetics & Developmental Biology | http://www.gradbiomed.pitt.edu/programs/molecular-genetics-development-biology |
| University of Pittsburgh | Molecular Pharmacology | http://www.gradbiomed.pitt.edu/programs/molecular-pharmacology |
| University of Pittsburgh | Neurobiology | https://www.cnup.pitt.edu/html/CNUP_Homepage.html |
| University of Rochester Medical Center | Biochemistry & Molecular Biology | https://www.urmc.rochester.edu/education/graduate/phd/biochemistry.aspx |
| University of Rochester Medical Center | Biophysics, Structural & Computational Biology | https://www.urmc.rochester.edu/education/graduate/phd/biophysics.aspx |
| University of Rochester Medical Center | Cell Biology of Disease | https://www.urmc.rochester.edu/education/graduate/phd/pathology.aspx |
| University of Rochester Medical Center | Cellular and Molecular Pharmacology and Physiology | https://www.urmc.rochester.edu/education/graduate/phd/pharmacology-and-physiology.aspx |
| University of Rochester Medical Center | Genetics, Development and Stem Cells | https://www.urmc.rochester.edu/education/graduate/phd/genetics.aspx |
| University of Rochester Medical Center | Immunology, Microbiology & Virology | https://www.urmc.rochester.edu/education/graduate/phd/immunology-microbiology-virology.aspx |
| University of Rochester Medical Center | Neuroscience | https://www.urmc.rochester.edu/education/graduate/phd/neurosciences.aspx |
| University of Rochester Medical Center | Toxicology | https://www.urmc.rochester.edu/education/graduate/phd/toxicology.aspx |
| University of Rochester Medical Center | Translational Biomedical Science | https://www.urmc.rochester.edu/education/graduate/phd/translational-biomedical-science.aspx |
| University of South Dakota School of Medicine | Basic Biomedical Sciences | https://www.usd.edu/medicine/basic-biomedical-sciences |
| University of Southern California | Biomedical and Biological Sciences | https://keck.usc.edu/pibbs/ |
| University of Southern California | Pharmaceutical and Translational Sciences | https://pharmgradprograms.usc.edu/phts/phd/admission/apply/ |
| University of South Carolina | Biological Sciences | https://www.sc.edu/study/colleges_schools/graduate_school/apply/degree_programs-application-requirements/biological-sciences-phd/index.php |
| University of Tennessee | Ecology and Evolutionary Biology | https://eeb.utk.edu/ |
| University of Tennessee | Microbiology | https://micro.utk.edu/ |
| UT Health San Antonio | Integrated Biomedical Sciences (IBMS) Program | https://www.uthscsa.edu/academics/biomedical-sciences/programs/integrated |
| UT Health San Antonio | Translational Science Program | https://www.uthscsa.edu/academics/biomedical-sciences/programs/translational-science-phd |
| University of Texas MD Anderson Cancer Center UTHealth | Graduate School of Biomedical Sciences | https://gsbs.uth.edu/ |
| University of Texas Southwestern Medical School | Graduate School of Biomedical Sciences (Umbrella PhD and MS) | http://www.utsouthwestern.edu/education/graduate-school/application-and-admissions/ |
| University of Utah | Bioscience Program (Umbrella PhD) | https://bioscience.utah.edu/#:~:text=The%20Bioscience%20PhD%20Programs%20are,at%20the%20University%20of%20Utah. |
| University of Utah | Molecular, Cellular, and Evolutionary Biology | https://www.biology.utah.edu/graduate/mceb/apply.php |
| University of Utah | Ecology, Evolution, and Organismal Biology | https://www.biology.utah.edu/graduate/eeob/apply.php |
| University of Vermont | Biology | https://www.uvm.edu/cas/biology/application-finances-support |
| University of Vermont | Cellular, Molecular, and Biomedical Sciences | https://www.uvm.edu/cmb |
| University of Vermont | Neuroscience | https://www.uvm.edu/neurosciencegrad |
| University of Virginia | Biomedical Sciences | http://bims.virginia.edu/ |
| University of Virginia | Biology | http://bio.as.virginia.edu |
| University of Virginia | Ecology and Evolutionary Biology | https://www.eebvirginia.org/ |
| University of Virginia | Psychology | https://psychology.as.virginia.edu/graduate-studies |
| University of Virginia | Biomedical Engineering | https://engineering.virginia.edu/departments/biomedical-engineering |
| University of Washington | Neuroscience | https://depts.washington.edu/neurogrd/prospective-students/admissions/ |
| University of Washington | Biochemistry | https://depts.washington.edu/biowww/ |
| University of Washington | Biology | https://www.biology.washington.edu/ |
| University of Washington | Chemical Engineering | https://www.cheme.washington.edu/ |
| University of Washington | Genome Sciences | https://www.gs.washington.edu/academics/gradprogram/index.htm |
| University of Washington | Immunology | http://www.immunology.washington.edu/graduate-program/admissions/admissions-faqs/ |
| University of Washington | Molecular Medicine and Mechanisms of Disease | http://www.pathology.washington.edu/academics/mmtp/phd-program/ |
| University of Washington | Molecular and Cellular Biology | https://www.washington.edu/students/gencat/academic/mcb.html |
| University of Wisconsin-Madison | Cell and Molecular Biology | https://cmb.wisc.edu/admissions/ |
| University of Wisconsin-Madison | Endocrinology | https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/ |
| University of Wisconsin-Madison | Integrated Program in Biochemistry | https://ipib.wisc.edu/ |
| University of Wisconsin-Madison | Microbiology | https://microbiology.wisc.edu/ps_apply.php |
| University of Wisconsin-Madison | Neuroscience | https://ntp.neuroscience.wisc.edu/ |
| University of Wisconsin-Madison | Nutritional Sciences | https://nutrisci.wisc.edu/graduate/ |
| University of Wisconsin-Madison | Pharmacology | https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/ |
| Van Andel Institute | Molecular and Cellular Biology | https://vaigs.vai.org/admissions/apply/ |
| Vanderbilt University | Biological Sciences | https://as.vanderbilt.edu/biosci/graduate/admission.php |
| Vanderbilt University | Interdisciplinary Graduate Program | https://medschool.vanderbilt.edu/igp/ |
| Vanderbilt University | Quantitative and Chemical Biology | https://medschool.vanderbilt.edu/qcb/ |
| Vanderbilt University | Neuroscience | https://medschool.vanderbilt.edu/brain-institute/new-vbi-homepage-inprogress/join-our-graduate-program/ |
| Virginia Commonwealth University | Biomedical Sciences Doctoral Portal | https://medschool.vcu.edu/education/bsdp/ |
| Virginia Commonwealth University | Clinical and Translational Research, Cancer and Molecular Medicine | http://bulletin.vcu.edu/graduate/office-research/center-clinical-translational-research/clinical-translational-sciences-phd-concentration-cancer-molecular-medicine/ |
| Virginia Tech | Biochemistry | https://www.biochem.vt.edu/ |
| Virginia Tech | Biological Sciences | https://www.biol.vt.edu/ |
| Wake Forest School of Medicine | Biomedical Graduate Programs (multiple programs) | https://school.wakehealth.edu/Education-and-Training/Graduate-Programs |
| Wake Forest School of Medicine | Molecular and Cellular Biosciences Track | https://school.wakehealth.edu/Education-and-Training/Graduate-Programs/Molecular-and-Cellular-Biosciences-Track#:~:text=The%20Molecular%20and%20Cellular%20Bioscience,to%20a%20specific%20research%20topic. |
| Washington State University | Biological Sciences | https://sbs.wsu.edu/ |
| Washington State University | Biological Systems Engineering | https://bsyse.wsu.edu/graduate-program/how-to-apply/ |
| Washington University in St. Louis | DBBS – Biochemistry, Biophysics & Structural Biology | http://dbbs.wustl.edu/divprograms/biophysics/Pages/BBSB.aspx |
| Washington University in St. Louis | DBBS – Developmental, Regenerative & Stem Cell Biology | http://www.dbbs.wustl.edu/divprograms/devbio/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Cancer Biology | http://dbbs.wustl.edu/divprograms/cancerbiology/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Computational & Systems Biology | http://dbbs.wustl.edu/divprograms/compbio/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Evolution, Ecology & Population Biology | http://www.dbbs.wustl.edu/divprograms/eepb/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Human & Statistical Genetics | http://dbbs.wustl.edu/divprograms/hsg/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Immunology | http://dbbs.wustl.edu/divprograms/immunology/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Molecular Genetics & Genomics | http://www.dbbs.wustl.edu/divprograms/genetics/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Molecular Microbiology & Microbial Pathogenesis | http://www.dbbs.wustl.edu/divprograms/micro/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Neurosciences | http://dbbs.wustl.edu/divprograms/neuro/Pages/default.aspx |
| Washington University in St. Louis | DBBS – Plant & Microbial Biosciences | http://dbbs.wustl.edu/divprograms/PlantMicroBioSci/Pages/default.aspx |
| Washington University in St. Louis | DDBS – Molecular Cell Biology | http://www.dbbs.wustl.edu/divprograms/cellbio/Pages/default.aspx |
| Wayne State University | Biochemistry, Immunology and Microbiology | https://biochemmicroimmuno.med.wayne.edu/ |
| Washington University in St. Louis | DDBS – Biomedical Informatics & Data Science | http://dbbs.wustl.edu/divprograms/BIDS/Pages/default.aspx |
| Weill Cornell Graduate School of Medical Sciences | BCMB Allied Program | https://gradschool.weill.cornell.edu/bcmb-allied |
| Weill Cornell Graduate School of Medical Sciences | Immunology & Microbial Pathogenesis Program | https://gradschool.weill.cornell.edu/programs/immunology-microbial-pathogenesis |
| Weill Cornell Graduate School of Medical Sciences | Neuroscience Program | https://gradschool.weill.cornell.edu/programs/neuroscience |
| Weill Cornell Graduate School of Medical Sciences | Pharmacology Program | https://gradschool.weill.cornell.edu/programs/pharmacology |
| Weill Cornell Graduate School of Medical Sciences | Physiology, Biophysics and Systems Biology Program | https://gradschool.weill.cornell.edu/programs/physiology-biophysics-systems-biology |
| West Virginia University | Biomedical Sciences | https://www.hsc.wvu.edu/resoff/graduate-education/phd-programs/biomedical-sciences/ |
| Wright State University | Biological Sciences (MS) | https://science-math.wright.edu/biology/master-of-science-in-biological-sciences |
| Yale University | Combined Program in Biological & Biomedical Sciences | https://medicine.yale.edu/bbs/ |
| Updated February 13, 2021 | ||
| Please email Joshua Hall ([email protected]) or tweet to @jdhallphd for additions or corrections to this list. |
